What are the latest Web 2.0 online apps programmed with?

So, what are the latest Web 2.0 applications using as their programming language?

I mean, twitter, pownce and all that good stuff. Are they developing everything with just HTML, CSS, PHP and MySQL or is there some other language in the mix?
Great responses so far…

If you could comment on the pros and cons of using Django or Ruby on Rails compared to LAMP, that would really help. Thanks!

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